statistical, or computational modeling. Proposals should be submitted to the core clusters (Ecosystem Sciences, Evolutionary Processes, Population and Community Ecology, and Systematics and Biodiversity Sciences). DEB also encourages interdisciplinary proposals that cross conceptual boundaries and integrate over levels of biological organization or across multiple spatial and temporal scales.Research addressing ecology and ecosystem science in the marine biome should be directed to the Biological Oceanography Program in the Division of Ocean Sciences; research addressing evolution and systematics in the marine biome should be directed to the Evolutionary Processes or Systematics and Biodiversity Science programs in DEB. All DEB programs also encourage proposals that leverage NSF-supported data networks, databases, centers, and other forms of scientific infrastructure, including but not limited to the National Ecological Observatory Network (NEON), Environmental Data Initiative (EDI), and Integrated Digitized Biocollections (iDigBio). Rules of Life Track proposals that integrate across the scales in biological sciences are solicited to support research that spans from the population, species, community and ecosystem scales normally funded by DEB, to organismal, cellular and molecular scales typically funded by other divisions in the Biological Sciences.This track provides new opportunities to advance our understanding of the Rules of Life by new mechanisms for review and funding of proposals that would not ordinarily fit well within one division in the Biological Sciences Directorate.